| 1 | Stems thinly arachnoid or glabrescent; stem wings narrow, spinose-lobed |
| 1' | Stems white-tomentose; stem wings conspicuous, lobulate, with numerous rigid spines |
| 2 | Leaves deeply pinnatisect, almost to midrib |
| 2' | Leaves irregularly and not very deeply lobed, incisions less than 50% in the middle of the leaf |
