R. D. Meikle, Flora of Cyprus 2. 1985
but much modified and supplemented
1Corolla yellow
1'Corolla white to purple
2Stems and fruits spiny
2'Stems and fruits not spiny
3Small shrubs with lanceolate, entire leaves
3'Herbs with ovate or oblong, lobed leaves
4Ripe fruit red or orange; upper parts of stems villose, with long, white, multicellular hairs
4'Ripe fruit black; upper parts of stems thinly clothed with short, incurved, subadpressed hairs